Clayton A Penniman, PhD
Professor & Department Chair
Biology
Office Information
Nicolaus Copernicus Hall
332
Biography

I am a marine biologist with specialization in seaweeds. I am also interested in watershed management and water quality of lower-order streams, particularly in developing and developed watersheds.

After getting my doctoral degree at the University of New Hampshire, I spent seven years at the UNH Jackson Estuarine Laboratory as a research scientist and then a Research Assistant Professor. Immediately prior to coming to CCSU, I spent two years as the Senior Scientist and Assistant Project Manager at US EPA/RI DEM Narragansett Bay Project.

I coordinate the Environmental Program in the Biology Department and teach a variety of courses in various aspects of aquatic ecology and environmental management (see below).

My current research interests are quite broad but focus on (1) water quality assessment of New England wadeable streams and rivers and (2) population ecology of selected Northwestern Atlantic seaweeds. I try to offer both undergraduate research projects and master’s thesis projects that are tailored to individual student interests within the scope of my laboratory’s research capabilities.

Publications, Research & Presentations

Recent student projects:

Water Quality Assessment and the Effect of Recreational Tubing on the Farmington River in New Hartford, Connecticut. Clark Strauss. Summer 2018.

Population Biology of the Red Seaweed, Chondrus crispus, in Long Island Sound. Marina Callegari and Patrick Regan. Spring 2018.

Water Quality Assessment of Kettle Brook in Windsor Locks, CT. Emily Dixon. Spring 2018.

Water Quality Assessment of Birge Pond,  Bristol, CT. Emily Brunet. Fall 2017.

Effects of Stormwater Runoff from the CCSU Campus on Water Quality in Sandy Brook. Ben Chagnot, Amanda Ross, Heather Bourgoin, and Magdalena Gwozdz. Fall 2017.

Water Quality Assessment of the Effects of a Water Pollution Control Facility on the Hockanum River in Vernon, Connecticut. Alexander Manos. Spring 2017.

Assessment of Water Quality in the Farmington River using Microbial Indicators of Fecal Pollution. Alyssa Roland. Spring 2016.

Water Quality Assessment of Selected Ponds in Terryville, Connecticut. Holly Geddes. Spring 2016.

Master’s theses:

Morris-Scata, Carol. L. 2017, Crayfish Dynamics in the Ten Mile River, Lebanon Connecticut. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 268 pp.

Weneck, Jillian. 2016. A Water Quality Assessment of Railroad Brook, Vernon, Connecticut. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 220 pp. 

Costello, Christopher. 2014. The Limnological Dynamics of a Small Urban Lake, Batterson Park Pond, Connecticut. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 311 pp.

King, Katelynn.  2014.  Microbial Water Quality Analysis of Bass and Piper Brooks in New Britain and Newington, Connecticut. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 170 pp.

Hall, Robyn. 2014. Chemical and Biological Water Quality Assessment of Clear Brook and the Nepaug River Tributaries to the Metropolitan District Commission’s (MDC) Nepaug Drinking Water Reservoir in New Hartford, CT. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  143 pp.

Bakaj, Magen.  2012.  Chemical Water Quality Assessment of the Southern Part of the Eightmile River in Lyme, Connecticut.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 244 pp.

Blessing, Kathleen.  2012.  Reproductive phenology of Polysiphonia subtilissima (Rhodophyta, Ceramiales).  Thesis, M.A. Biology, Department of Biology, CCSU.  47 pp.

Imfeld, Isabel T.  2012.  Effect of a Dog Park on Water Quality in an Adjacent River.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU. 65 pp.

Marrett, Marva.  2012.  Microbiological Water Quality of the Southern Part of the Eightmile River Watershed - A Baseline Study. Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  166 pp.

Mozejko, Agata.  2012.  Microbial Water Quality Assessment of the East Branch of the Eightmile River in Lyme, East Haddam and Salem, Connecticut.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  105 pp.

O’Dea, Daniel J. 2012.  Examining the fossil assemblage of a presumed CO2-sequestering lake (the Grube Messel lagerstatte, Middle Eocene), with a view to explaining its origin and composition.  Thesis, M.S. in Biology.  Department of Biology, CCSU.  119 pp.

Koffman, Nancy.  2011.  Disturbance and the Gulf Coast Environment: an investigation of hurricane effects on oil spills and coastal ecology.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  64 pp.

Protopopova, Daria L. 2011.  Chemical defenses in the Class Phaeophyceae for protection from marine herbivores.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  51 pp.

Sarkar, Babul Kanti.  2011.  Leaf Litter Processing by Stream Macroinvertebrates.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  61 pp.      

Tipton, Michelle.  2010.  Chemical Quality Assessment of the East Branch of the Eight Mile River in Lyme/Salem, CT.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  246 pp.

Grabarz, Theodore.  2007.  Assessment of Rocky Intertidal Habitat on Outer Island, using Paired Transects Experiencing Different Wave Exposures.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  137 pp.

Henry, Lidia A.  2006.  Aquatic Plants of Outer Island.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.

Joaquin, Mary Elizabeth.  2006.  A Chemical Water Quality Analysis of the Eightmile River, West Branch.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  234 pp.

LaFountain, Amy.  2006.  A Baseline Microbiological Water Quality Study of the Western

Subwatershed of the Eightmile River.  Thesis, M.A. in Biology: Ecology and Environmental Sciences.  Department of Biology, CCSU.  130 pp.

Dopirak, William J., Jr.  2002.  Analyses of Competing Macrophytes in a Eutrophic Lake and in Microcosm Experiments.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.

Dworetzky, Barbara L.  2000.  The Impact of Combined Sewer Overflows on the Water Quality of Wethersfield Cove, Wethersfield, CT  March-August 1999.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.  107 pp. and appendices.

Dunn, Joseph G.  1998.  Comparative Analysis of the Benthic Infauna and Sediment for Two Small Estuaries in Connecticut.  Thesis, M.S. Biology, Department of Biological Sciences,  CCSU.  64 pp.

Consolati (Suljak), Kristen L.  1997.  Water Quality Assessment of the Mattabesset River During High and Low Flow Periods.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.  48 pp. and appendices.

Costa, Mary Ann.  1997.  A Watershed-Based, Water-Quality Study of the Lieutenant River in Central Coastal Connecticut.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.  76 pp. and appendices.

Kruse, Valerie A.  1997.  Water Quality Monitoring for the Black Hall River, Old Lyme, Connecticut.  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.  33 pp. and appendices.

Tata, Joanna V.  1996.  Investigations of Isozyme Variability within Long Island Sound Populations of Eelgrass (Zostera marina).  Thesis, M.A. Biology, Department of Biological Sciences, CCSU.  61 pp. and appendices.

Publications:

Tessier, J. and C. Penniman.  2006.  An inquiry-based laboratory design for microbial ecology.  Bioscence 32(4):6-11.

Penniman, C.A.  2001.  Regional Effects of Air Pollution (2001).  Web-based Regional

Supplement to: Nebel, B.J. and R.T. Wright.  2001.  Environmental Science: Toward a Sustainable Future  Eighth Edition.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A.  2001.  Cleaning Up Boston Harbor (2001).  Web-based Regional

Supplement to: Nebel, B.J. and R.T. Wright.  2001.  Environmental Science: Toward a Sustainable Future  Eighth Edition.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A.  2001.  Atmospheric Fertilization of Estuaries - Watersheds and Airsheds (2001).  Web-based Regional Supplement to: Nebel, B.J. and R.T. Wright.  2001.  Environmental Science: Toward a Sustainable Future  Eighth Edition.  Prentice-Hall, Inc.,  Upper Saddle River, NJ.

Penniman, C.A.  2001.  The Northern Forest (2001).  Web-based Regional Supplement to: Nebel, B.J. and R.T. Wright.  2001.  Environmental Science: Toward a Sustainable Future  Eighth Edition.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A.  2001.  Suburban Sprawl in the Northeast (2001).  Web-based Regional Supplement to: Nebel, B.J. and R.T. Wright.  2001.  Environmental Science: Toward a Sustainable Future  Eighth Edition.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A. 1999. Regional Effects of Air Pollution. [www.prenhall.com/divisions/esm/app/ nebel/html/updates/update18.html] WWW supplement to Nebel, B.J. and R.T. Wright. 1999.  Environmental ScienceSixth Ed.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A. 1999. Cleaning up Boston Harbor. [www.prenhall.com/divisions/esm/app/ nebel/html/updates/update19.html] WWW supplement to Nebel, B.J. and R.T. Wright. 1999.  Environmental ScienceSixth Ed.  Prentice-Hall, Inc., Upper Saddle River, NJ.

Penniman, C.A. 1999. Atmospheric Fertilization of Estuaries - Watersheds and Airsheds. [www.prenhall.com/divisions/esm/app/nebel/html/updates/update20.html] WWW supplement to Nebel, B.J. and R.T. Wright. 1999.  Environmental ScienceSixth Ed.  Prentice-Hall, Inc., Upper Saddle Rive, NJ.

Penniman, C.A. 1999. The Northern Forest. [www.prenhall.com/divisions/esm/app/ nebel/html/updates/update20.html] WWW supplement to Nebel, B.J. and R.T. Wright. 1999.  Environmental ScienceSixth Ed.  Prentice-Hall, Inc., Upper Saddle Rive, NJ.

Penniman, C.A. 1999. National Estuarine Research Reserves in the northeast -  Research, Education, and Management. [www.prenhall.com/divisions/esm/app/nebel/html/updates/ update21.html] WWW supplement to Nebel, B.J. and R.T. Wright. 1999.  Environmental ScienceSixth Ed.  Prentice-Hall, Inc., Upper Saddle Rive, NJ.

Penniman, C.A., M.A. Costa, and E.C. Carruthers.  1998.  An On-line Intertidal Transect at Outer Island, Long Island Sound, Connecticut [www.biology.ccsu.edu/intertidal/main.htm].

Penniman, C.A. and C. Yarish.  1992.  Effects on Nutrient Enrichment on Kelp (Laminaria longicruris) from Long Island Sound.  In: Proceedings of the Long Island Sound Research Conference, 23-24 October 1992.  Univ. Connecticut Sea Grant College Program, Avery Point, CT.

Narragansett Bay Project*.  1992.  Comprehensive Conservation and Management Plan for Narragansett Nay  Final Report, December 1992.  RI Department of Environmental Management, Providence, RI.  500 pp.  [* Final version released after approval by EPA Administrator William Reilly and RI Governor Bruce Sundlun; I prepared sections of final draft and submitted final draft for approval before leaving the Narragansett Bay Project.]

Penniman, C.A.  1992.  Management of Living Marine Resources in Narragansett Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.

Penniman, C.A., ed.  1992.  Science and the Narragansett Bay Comprehensive Conservation and Management Plan.  A Symposium Sponsored by the Narragansett Project, Rhode Island Sea Grant Program and University of Rhode Island Coastal Institute, 17 February 1992, Narragansett, RI.  Book of Abstracts, 73 pp.

Penniman, C.A., C.A. Karp, L. Remington and R.R. Zingarelli.  1991.  Control of Nutrient Inputs to Narragansett Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  122 pp.

Penniman, C.A., R.R. Zingarelli, C.A. Karp, K. Kipp and L. Remington.  1991.  Control of Toxics Inputs to Narragansett Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  59 pp.

Dixon, A.M., J. Myers, C. Penniman and R. Ribb.  1991.  Critical Resource Area Protection, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  41 pp.

Penniman, C.A.  1991.  Water quality effects of water quality standard enforcement: Industrial pretreatment in Rhode Island.  pp. 183-190, in: G.H. Flock, ed.  Water Quality Standards for the 21st Century, Proceedings of a Conference, 10-12 December, US EPA, Arlington, Virginia.  US EPA, Office of Water, Washington, DC.

Mathieson, A.C. and C.A. Penniman.  1991.  Floristic patterns and numerical classification of New England estuarine and open coastal seaweed populations.  Nova Hedwigia 52:453-485.

Monahan, R., S. Beede, J. Costa, K. Kipp, C. Penniman, C. Pring-Ham, B. Rosinoff and J. Sulak.  1991.  Controlling nitrogen loading to coastal waters.  pp. 3068-3080 in: O.T. Magoon et al., editors.  Proceedings of the Seventh Symposium on Coastal and Ocean Management, 8-12 July 1991, Long Beach, CA. 

Grizzle, R.E. and C.A. Penniman.  1991.  Sediment profile imaging data and benthic macrofaunal community changes along an organic enrichment gradient in a coastal lagoon.  Mar. Ecol. Prog. Ser. 74:249-261.

Mathieson, A.C., C.A. Penniman and L.G. Harris.  1991.  Northwest Atlantic Rocky Shore Ecology.  pp. 109-191 in: P. Nienhuis and A.C. Mathieson, eds.  Ecosystems of the World  Intertidal and Littoral Ecosystems.  Elsevier Scientific Publishing Company, Amsterdam. 

Yarish, C., C.A. Penniman and B. Egan.  1990.  Reproductive responses of Laminaria longicruris de la Pyl. to nutrient enrichment.  Proc. Int. Seaweed Symp. 13.  Hydrobiologia 204/205: 505-511.

Karp, C.A. and C.A. Penniman.  1990.  Boater Waste, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  23 pp.

Penniman, C.A., C.A. Karp and S. Hale.  1990.  Long-term Monitoring in Narragansett Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  36 pp.

Dixon, A., C.A. Karp and C.A. Penniman.  1990.  Mount Hope Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  23 pp.

Karp, C.A., C.A. Penniman, R. Zingarelli and A. Dixon.  1990.  Sewage contamination (Pathogens) of Narragansett Bay, Briefing Paper for the Narragansett Bay Project Comprehensive Conservation and Management Plan.  Narragansett Bay Project, Providence, RI.  54 pp.

Penniman, C.A. and L.G. Ward.  1990.  Little Bay Bathymetric Chart.  Tech. Rep. NH Coastal Program, NH Office of State Planning, Concord.  6 pp. + charts.

Penniman, C.A.  1990.  Summary of a panel discussion on the future of seaweed utilization and aquaculture.  pp. 151-158, in: C. Yarish, C.A. Penniman and P. Van Patten (editors).  Proceedings of the Symposium on Economically Important Marine Plants of the Atlantic: Their Biology and Cultivation.  Symposium at the University of Connecticut, Avery Point, Connecticut, October 3-4, 1988.  Univ. Conn. Sea Grant College Program, Avery Point, CT.

Yarish, C., C.A. Penniman and P. Van Patten (editors).  1990.  Proceedings of the Symposium on Economically Important Marine Plants of the Atlantic: Their Biology and Cultivation.  Symposium at the University of Connecticut, Avery Point, Connecticut, October 3-4, 1988.  Univ. Conn. Sea Grant College Program, Avery Point, CT.  158 pp.

Penniman, C.A., L.G. Ward, R.E. Grizzle and F.E. Anderson.  1989.  Great Bay Bathymetric Chart.  Tech. Rep. NH Coastal Program, NH Office of State Planning, Concord.  4 pp. + charts.

NH Office of State Planning.  1989.  Environmental Description of the Great Bay Estuary and the Research Management Plan for the Great Bay Estuarine Research Reserve.  NH Office of State Planning, Concord. 

Penniman, C.A.  1989.  Water quality in the Great Bay Estuary.  pp. 2-3, in: New Hampshire Cooperative Extension Water News, issue 6 (reprinted from The Great Bay Newsletter, Winter 1989), Durham, NH.

Penniman, C.A, F.T. Short and S.A. Adams.  1989.  Bibliography of research on the Great Bay Estuary and adjacent upland region.  Jackson Estuarine Laboratory Contribution number 220 and University of New Hampshire Sea Grant College Program Publication UNHMP-AR-SG-89-2.  113 pp.

Penniman, C.A. and A.C. Mathieson.  1987.  Variation in chemical composition of Gracilaria tikvahiae McLachlan (Gigartinales, Rhodophyta) in the Great Bay Estuary, New Hampshire.  Bot. Mar. 30:525-534.

Mathieson, A.C. and C.A. Penniman.  1986.  A phytogeographic interpretation of the marine flora from the Isles of Shoals, U.S.A.  Bot. Mar. 29:413-434.

Mathieson, A.C. and C.A. Penniman.  1986.  Species composition and seasonality of New England seaweeds along an open coastal - estuarine gradient.  Bot. Mar. 29:161-176.

Penniman, C.A., A.C. Mathieson and C. Emerich Penniman.  1986.  Reproductive phenology and growth of Gracilaria tikvahiae McLachlan (Gigartinales, Rhodophyta) in the Great Bay Estuary, New Hampshire.  Bot. Mar. 29:147-154.

Penniman, C.A. and A.C. Mathieson.  1985.  Photosynthesis of Gracilaria tikvahiae McLachlan (Gigartinales, Rhodophyta) from the Great Bay Estuary, New Hampshire. Bot. Mar. 28:427-435.

Mathieson, A.C., C.A. Penniman, P.K. Busse and E.T. Gallagher.  1982.  The effects of ice on Ascophyllum nodosum within the Great Bay Estuary System of N.H.-Maine.  J. Phycol. 18:331-336.

Courses Taught

BIO 100 Search In Biology: Introduction to Marine Science (Summer, online)

BIO 132 Introductory Ecology (Summer, online)

BIO 133 Laboratory in Introductory Ecology (Summer)

BIO 150 Introductory Ecology: Long Island Sound (Summer)

BIO 315 Microbial Ecology (Spring semesters and Summer)

BIO 407 Stream Ecology (Fall semesters)

BIO 425 Biology of Marine and Freshwater Algae (Fall semesters, Summer)

BIO 434 Ecology of Inland Waters (Fall semesters)

BIO 436 Environmental Resources and Management (Spring semesters)

BIO 438 Aquatic Pollution (Spring semesters)

BIO 507 Advanced Stream Ecology (Fall semesters)

BIO 508 Coastal Ecology (Summer)

BIO 509 Laboratory in Coastal Ecology (Summer)

BIO 515 Foundations in Ecology (Summer, online)

BIO 516 Advanced Biology of Marine and Freshwater Algae (Fall semesters, Summer)

BIOI 538 Advanced Aquatic Pollution (Spring semesters)